| On the Origin of Species, 6th Edition | 1 |
| An Historical Sketch Of The Progress Of Opinion On The Origin Of Species, Previously To The Publication Of The First Edition Of This Work | 2 |
| Introduction | 3 |
| Chapter 1 Variation Under Domestication | 4 |
| 1. Causes of Variability | 5 |
| 2. Effects Of Habit And Of The Use Or Disuse Of Parts; Correlated Variation; Inheritance | 6 |
| 3. Character Of Domestic Varieties; Difficulty Of Distinguishing Between Varieties And Species; Origin Of Domestic Varieties From One Or More Species | 7 |
| 4. Breeds Of The Domestic Pigeon, Their Differences And Origin | 8 |
| 5. Principles Of Selection Anciently Followed, And Their Effects | 9 |
| 6. Unconscious Selection | 10 |
| 7. Circumstances Favourable To Man's Power Of Selection | 11 |
| Chapter 2 Variation Under Nature | 12 |
| 1. | 13 |
| 2. Individual Differences | 14 |
| 3. Doubtful Species | 15 |
| 4. Wide-Ranging, Much Diffused, And Common Species Vary Most | 16 |
| 5. Species Of The Larger Genera In Each Country Vary More Frequently Than The Species Of The Smaller Genera | 17 |
| 6. Many Of The Species Included Within The Larger Genera Resemble Varieties In Being Very Closely, But Unequally, Related To Each Other, And In Having Restricted Ranges | 18 |
| 7. Summary | 19 |
| Chapter 3 Struggle For Existence | 20 |
| 1. | 21 |
| 2. The Term, Struggle For Existence, Used In A Large Sense | 22 |
| 3. Geometrical Ratio Of Increase | 23 |
| 4. Nature Of The Checks To Increase | 24 |
| 5. Complex Relations Of All Animals And Plants To Each Other In The Struggle For Existence | 25 |
| 6. Struggle For Life Most Severe Between Individuals And Varieties Of The Same Species | 26 |
| Chapter 4 Natural Selection; Or The Survival Of The Fittest | 27 |
| 1. | 28 |
| 2. Sexual Selection | 29 |
| 3. Illustrations Of The Action Of Natural Selection, Or The Survival Of The Fittest | 30 |
| 4. On The Intercrossing Of Individuals | 31 |
| 5. Circumstances Favourable For The Production Of New Forms Through Natural Selection | 32 |
| 6. Extinction Caused By Natural Selection | 33 |
| 7. Divergence Of Character | 34 |
| 8. The Probable Effects Of The Action Of Natural Selection Through Divergence Of Character And Extinction, On The Descendants Of A Common Ancestor | 35 |
| 9. On The Degree To Which Organisation Tends To Advance | 36 |
| 10. Convergence Of Character | 37 |
| 11. Summary Of Chapter | 38 |
| Chapter 5 Laws Of Variation | 39 |
| 1. | 40 |
| 2. Effects Of The Increased Use And Disuse Of Parts, As Controlled By Natural Selection | 41 |
| 3. Acclimatisation | 42 |
| 4. Correlated Variation | 43 |
| 5. Compensation And Economy Of Growth | 44 |
| 6. Multiple, Rudimentary, And Lowly-Organised Structures Are Variable | 45 |
| 7. A Part Developed In Any Species In An Extraordinary Degree Or Manner, In Comparison With The Same Part In Allied Species, Tends To Be Highly Variable | 46 |
| 8. Specific Characters More Variable Than Generic Characters | 47 |
| 9. Secondary Sexual Characters Variable | 48 |
| 10. Distinct Species Present Analogous Variations, So That A Variety Of One Species Often Assumes A Character Proper To An Allied Species, Or Reverts To Some Of The Characters Of An Early Progenitor | 49 |
| 11. Summary | 50 |
| Chapter 6 Difficulties Of The Theory | 51 |
| 1. | 52 |
| 2. On The Absence Or Rarity Of Transitional Varieties | 53 |
| 3. On The Origin And Transition Of Organic Beings With Peculiar Habits And Structure | 54 |
| 4. Organs Of Extreme Perfection And Complication | 55 |
| 5. Modes Of Transition | 56 |
| 6. Special Difficulties Of The Theory Of Natural Selection | 57 |
| 7. Organs Of Little Apparent Importance, As Affected By Natural Selection | 58 |
| 8. Utilitarian Doctrine, How Far True: Beauty, How Acquired | 59 |
| 9. Summary: The Law Of Unity Of Type And Of The Conditions Of Existence Embraced By The Theory Of Natural Selection | 60 |
| Chapter 7 Miscellaneous Objections To The Theory Of Natural Selection | 61 |
| Chapter 8 Instinct | 62 |
| 1. | 63 |
| 2. Inherited Changes Of Habit Or Instinct In Domesticated Animals | 64 |
| 3. Special Instincts | 65 |
| 4. Instincts Of The Cuckoo | 66 |
| 5. Slave-Making Instinct | 67 |
| 6. Cell-Making Instinct Of The Hive-Bee | 68 |
| 7. Objections To The Theory Of Natural Selection As Applied To Instincts: Neuter And Sterile Insects | 69 |
| 8. Summary | 70 |
| Chapter 9 Hybridism | 71 |
| 1. | 72 |
| 2. Degrees Of Sterility | 73 |
| 3. Laws Governing The Sterility Of First Crosses And Of Hybrids | 74 |
| 4. Origin And Causes Of The Sterility Of First Crosses And Of Hybrids | 75 |
| 5. Reciprocal Dimorphism And Trimorphism | 76 |
| 6. Fertility Of Varieties When Crossed, And Of Their Mongrel Offspring, Not Universal | 77 |
| 7. Hybrids And Mongrels Compared, Independently Of Their Fertility | 78 |
| 8. Summary Of Chapter | 79 |
| Chapter 10 On The Imperfection Of The Geological Record | 80 |
| 1. | 81 |
| 2. On The Lapse Of Time, As Inferred From The Rate Of Deposition And Extent Of Denudation | 82 |
| 3. On The Poorness Of Palaeontological Collections | 83 |
| 4. On The Absence Of Numerous Intermediate Varieties In Any Single Formation | 84 |
| 5. On The Sudden Appearance Of Whole Groups Of Allied Species | 85 |
| 6. On The Sudden Appearance Of Groups Of Allied Species In The Lowest Known Fossiliferous Strata | 86 |
| Chapter 11 On The Geological Succession Of Organic Beings | 87 |
| 1. | 88 |
| 2. On Extinction | 89 |
| 3. On The Forms Of Life Changing Almost Simultaneously Throughout The World | 90 |
| 4. On The Affinities Of Extinct Species To Each Other, And To Living Forms | 91 |
| 5. On The State Of Development Of Ancient Compared With Living Forms | 92 |
| 6. On The Succession Of The Same Types Within The Same Areas, During The Later Tertiary Periods | 93 |
| 7. Summary Of The Preceding And Present Chapters | 94 |
| Chapter 12 Geographical Distribution | 95 |
| 1. | 96 |
| 2. Single Centres Of Supposed Creation | 97 |
| 3. Means Of Dispersal | 98 |
| 4. Dispersal During The Glacial Period | 99 |
| 5. Alternate Glacial Periods In The North And South | 100 |
| Chapter 13 Geographical Distribution--continued | 101 |
| 1. Fresh-Water Productions | 102 |
| 2. On The Inhabitants Of Oceanic Islands | 103 |
| 3. Absence Of Batrachians And Terrestrial Mammals On Oceanic Islands | 104 |
| 4. On The Relations Of The Inhabitants Of Islands To Those Of The Nearest Mainland | 105 |
| 5. Summary Of The Last And Present Chapters | 106 |
| Chapter 14 Mutual Affinities Of Organic Beings: Morphology -- Embryology -- Rudimentary Organs | 107 |
| 1. Classification | 108 |
| 2. Analogical Resemblances | 109 |
| 3. On The Nature Of The Affinities Connecting Organic Beings | 110 |
| 4. Morphology | 111 |
| 5. Development And Embryology | 112 |
| 6. Rudimentary, Atrophied, And Aborted Organs | 113 |
| 7. Summary | 114 |
| Chapter 15 Recapitulation And Conclusion | 115 |
| Glossary Of The Principal Scientific Terms Used In The Present Volume | 116 |
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